Fitness Theory Practice Exam – Body BlueprintForgot password? Don't have an account? The basis of inclusive fitness theory is Hamilton's rule, which specifies the conditions for the spread of a gene for any of four types of social action, namely cooperation, altruism, selfishness, and spite. Hamilton's rule predicts that cooperation and selfishness can evolve at all levels of relatedness, whereas altruism requires positive relatedness and spite requires negative relatedness. When groups are non-clonal, within-group conflict arises because not all group members have identical inclusive fitness optima.
Fitness Theory Practice Exam
ONE of the central dogmas of modern behavioral ecology is that blood kinship plays a critical role in understanding the evolution of social behavior, particularly of costly social behavior such as altruism and cooperation. But it was not always so, and what I would like to do in this Perspectives is provide some historical context that led up to William Hamilton's seminal work developing inclusive fitness theory. The story begins, not surprisingly, with Charles Darwin. The worker bees that sacrifice themselves to protect their hives—the ultimate example of animal altruism—were deeply troubling to Darwin. If increased reproduction is the currency of natural selection, then altruists should disappear—and fast. Eventually, however, Darwin came up with an explanation. Since sterile worker bees were helping their blood relatives—especially the queen—Darwin hypothesized that natural selection might favor altruism at the level of blood kin.
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